sound production (as in American Woodcock) or muffling (as in owls)
hearing
protection
cleanliness
water repellency
water transport (e.g., Desert Sand grouse)
tactile sensation
support (e.g., tail feathers of a woodpecker)
display
Feathers host bacteria and ectoparasites. They require preening. They molt periodically.
Interesting feather facts:
The red on male house finches comes from carotenoid pigments generated by they eat. Females prefer more brightly colored males, perhaps because it indicates better foraging capability.
Dark-colored feathers have more melanin in them, and are less likely to fray.
The blue in bluebird feathers is produced by air-filled cavities within the keratin of the cells in the feathers. They are not iridescent like the feathers of a hummingbird, and thus the hue looks the same regardless of the viewing angle. The feathers of a male bluebird reflect a lot of UV light which makes them brighter. Feathers of females look more subdued because their structure is different. Humans can't see UV wavelengths, but birds can.
Brighter male bluebirds tend to pair earlier, feed young more often and fledge more young than males with weaker ultraviolet sheens (Sifferman and Hill, 2003)
Most birds have thousands of feathers - songbirds typically have 2000-4000, 30-40% of which are on the head and neck.
Tiny muscles raise and lower feathers and twist them or pull them closer together.
Preening keeps feathers moist and flexible, and maintains waterproofing and insulation capabilities, and protects them against bacteria and fungi that digest the keratin in the feathers, and may discourage feather lice.
A 44 million year old fossilized bird louse was found with feathers in its gut (Wappler et al 2004). As many as 12 species of lice may be living on the feathers of one bird. There are more than 150 species total, including chewing lice, louse flies and feather mites. When birds scratch their heads, they are probably delousing.
Adult birds typically molt after breeding, replacing all their plumage. Feathers can change appearance as a result of wear. Starlings have spotted plumage in winter, and then as the feather tips wear off, the look changes to sleek and glossy. Some birds undergo partial molts in addition to the main annual molt.
In some Passerine birds, males do not have full adult breeding plumage until after their first potential breeding season.
The feather coat on a dead bird typically weights 2-3 x as much as the skeleton.
Some birds use feathers, probably for insulation, in the nest cup.